ELNAIS SPECIES

The following table includes information (Species name, First Sighting in Greece, Pathway, Status, Establishment success) of the non-indigenous species recorded in Greece in the database provided by the hELlenic Network on AquatIc Species (ELNAIS). The categories of pathways of introduction of introductions are the following: TS = Transport-Stowaway (vessels), UNA = Unaided,
REL = Releases, UNK = Unknown, TC = Transport-Contaminant, EC = Escape from Confinemed.

SPECIES NAMEFIRST SIGHTINGPATHWAY NEW/CBDSTATUSESTABLISHMENT SUCCESS
Cercozoa, Endomyxa
Marteilia refringens Cavalier-Smith, 20021997TCALest
Marinomyxa marina (Lipkin & Avidor) Kolátková, Čepička, Hoffman et Vohník, 20202018TCALest
Foraminifera
Amphistegina lobifera Larsen, 19761955-64UNAALinv
Amphistegina lessonii d'Orbigny in Guérin-Méneville, 18321974UNAALest
Clavulina cf. multicamerata Chapman, 19072012TSALcas
Heterostegina depressa d’Orbigny, 1826<1988UNA, TSALest
Triloculina cf. fichteliana d'Orbigny, 18392006UNA, TSALest
Planogypsina acervalis (Brady, 1884)1909UNAALest
Sigmamiliolinella australis (Parr, 1932)2001UNALest
Ochrophyta
Colpomenia peregrina Sauvageau1986TS-hullsALest
Cutleria multifida (Turner) Greville1950TS-Angl/fisALest
Dictyota cyanoloma Tronholm, De Clerck, A.Gómez-Garreta & Rull Lluch2013TS-hullsALcas
Pseudochattonella verruculosa (Hara & Chihara) Tanabe-Hosoi, Honda, Fukaya, Inagaki & Sako1998TSALcas
Pylaiella littoralis (Linnaeus) Kjellman1967TS-hullsALque
Scytosiphon dotyi Wynne2013TS-hullsALque
Stypopodium schimperi (Kützing) M.Verlaque & Boudouresque1996UNAALinv
PlantaeRhodophyta
Antithamnion amphigeneum A.Millar, 19902013ALunk
Asparagopsis armata Harvey2010UNAALcas
Asparagopsis taxiformis (Delile) Trevisan de Saint-Léon1992TS-Angl/fisALinv
Botryocladia madagascariensis G. Feldmann2006TS-Angl/fisALest
Ceramium strobiliforme Lawson & John2001TS-hullsALque
Chondria collinsiana Howe1980TS-hullsALque
Colaconema codicola (Børgesen) Stegenga, J.J.Bolton & R.J.Anderson,2019ALunk
Hypnea anastomosans Papenfuss, Lipkin & P.C. Silva2008UNAALest
Hypnea cornuta (Kützing) J. Agardh1894TS-Angl/fisALest
Hypnea spinella (C.Agardh) Kützing1979TS-Angl/fisALest
Hypnea valentiae (Turner) Montagne2009TS-hullsALcas
Laurencia caduciramulosa Masuda & Kawaguchi2001TS-hullsALest
Lophocladia lallemandii (Montagne) F. Schmitz1908TS-Angl/fisALest
Melanothamnus harveyi (Bailey) Díaz-Tapia & Maggs2006TS-hullsALest
Sarconema scinaioides Børgesen1980ECALcas
Womersleyella setacea (Hollenberg) R.E. Norris1988TS-Angl/fisALinv
Chlorophyta
Caulerpa cylindracea Sonder1993UNAALinv
Caulerpa racemosa var. lamourouxii f. requienii (Montagne) Weber van Bose1956UNAALest
Caulerpa taxifolia var. distichophylla (Sonder) Verlaque, Huisman & Procaccini2010UNAALest
Codium fragile (Suringar) Hariot1992TS-Angl/fisALinv
Tracheophyta
Halophila stipulacea (Forsskål) Ascherson1894TS, UNAALest
AnimaliaPorifera
Paraleucilla magna Klautau, Monteiro & Borojevic, 20042014TCALest
Ctenophora
Beroe ovata Mayer, 19122004UNAALcas
Mnemiopsis leidyi (Agassiz, 1865)1990UNAALest
Cnidaria
Hydrozoa
Clytia linearis (Thornely, 1900)1977UNAALest
Sertularia marginata (Kirchenpauer, 1864)1990TSALest
Scyphozoa
Cassiopea andromeda (Forsskål, 1775)1942UNAALest
Phylloriza punctata von Lendenfeld, 18842005UNALunk
Rhopilema nomadica Galil, 19902006UNAALcas
Chrysaora cf. achlyos Martin, Gershwin, Burnett, Cargo & Bloom, 19972018TSALcas
Platyhelminthes
Glyphidohaptor plectocirra (Paperna, 1972)2010TCALest
Mollusca
Bivalvia
Anadara transversa (Say, 1822)1993TS-hullsALest
Brachidontes pharaonis (P. Fischer, 1870)1975UNA, TSALest
Chama asperella Lamarck, 18192007UNA, TSALest
Chama pacifica Broderip, 18352005UNAALest
Clementia papyracea (Gmelin, 1791)1985UNAALcas
Crassostrea/Magallana sp./spp.≤1989EC?ALest
Dendostrea cf. folium (Linnaeus, 1758)2005UNA, TSALinv
Fulvia fragilis (Forsskål in Niebuhr, 1775)1997UNA, TSALinv
Isognomon legumen (Gmelin, 1791)2016UNAALest
Isognomon aff. australica (Reeve, 1858)2016TS, CORALunk
Isognomon bicolor (C.B. Adams, 1845)2022ALunk
Malleus regula (Forsskål in Niebuhr, 1775) ≤1996UNAALest
Mya arenaria Linnaeus, 17581984TS, TCALest
Petricolaria pholadiformis Lamarck, 18181985TSALest
Pinctada imbricata Röding, 17981961EC, TS-hullsALinv
Septifer cumingii (Dunker, 1855)2010UNAALest
Spondylus cf. spinosus Schreibers, 17932008-13TS-hullsALcas
Gastropoda
Atys angustatus E. A. Smith, 18722017UNAALcas
Atys ehrenbergi (Issel, 1869)2016CORALunk
Acteocina mucronata (Philippi, 1849)1991UNAALcas
Baeolidia moebii Bergh, 18882021ALest
Biuve fulvipunctata (Baba, 1938)??ALunk
Bulla arabica Malaquias & Reid, 20081998TSALest
Bursatella leachii (De Blainville, 1817)1975UNA, TSALest
Cerithidium perparvulum (Watson, 1886)2010UNAALcas
Cerithiopsis pulvis (A. Issel, 1869)2010UNAALest
Cerithiopsis tenthrenois (Melvill, 1896)1994UNAALcas
Cerithium scabridum Philippi, 18482007UNAALest
Cingulina isseli (Tryon, 1886)2021TSALest?
Conomurex persicus (Swainson, 1821)1983UNA, TSALinv
Coryphellina rubrolineata O'Donoghue, 19292009UNAALest
Crepidula fornicata (Linnaeus, 1758)1985TC, TSALest
Cycloscala hyalina (G.B. Sowerby II, 1844)<2019UNAALest
Dikoleps micalii Agamennone, Sbrana, Nardi, Siragusa & Germanà, 20202016COR? UNALcas
Diodora funiculata (Reeve, 1850) 2013TSALest
Elysia nealae Ostergaard, 19552019unkALcas
Ergalatax junionae Houart, 20082007UNAALest
Eunaticina papilla (Gmelin, 1791)2020TSALcas/est?
Euthymella colzumensis (Jousseaume, 1898)2017TS/COR?ALest
Goniobranchus annulatus (Eliot, 1904)2004UNA, TSALest
Lamprohaminoea ovalis (Pease, 1868)2001UNA, TSALest
Heliacus implexus (Mighels, 1845)2019ALcas
Hypselodoris infucata (Rüppell & Leuckart, 1831)2007UNAALest
Melanella orientalis Agamennone, Micali & Siragusa, 20202016COR? UNALest?
Melibe viridis (Kelaart, 1858)1970UNA, TSALest
Mnestia girardi (Audouin, 1826)1994UNAALest
Nerita sanguinolenta Menke, 18291969UNAALcas
Oscilla galilae Bogi, Karhan & Yokeş, 20122016UNAALcas
Plocamopherus ocellatus Rüppell & Leuckart, 18282020UNAALcas
Polycerella emertoni Verrill, 18811995TSALcas
Pyrunculus fourierii (Audouin, 1826)2013UNAALcas
Pyrgulina pupaeformis (Souverbie, 1865)2017TSALest
Rapana venosa (Valenciennes, 1846)1986UNAALcas
Rhinoclavis kochi (Philippi, 1848)2016UNAALest
Ringicula minuta H. Adams, 18722019TSALcas
Smaragdia souverbiana (Montrouzier, 1863)≤1993UNAALest
Sticteulima sp. [cf. lentiginosa] (A. Adams, 1861)2015UNA, TSALcas
Sinezona plicata (Hedley, 1899)2019TSALcas
Syphonota geographica (A. Adams & Reeve, 1850)2002UNAALest
Syrnola fasciata Jickeli, 18822012UNAALest
Viriola sp.[cf. bayani] Jousseaume, 18842016UNAALest
Cephalopoda
Sepioteuthis lessoniana Férussac in Lesson, 1831 complex 2009UNACRYest
Annelida
Axionice medusa (Savigny in Lamarck, 1818)1976UNAALest
Branchiomma bairdi (McIntosh, 1885)2014TS-hullsALest
Branchiomma luctuosum (Grube, 1869)1989TS-hullsALest
Caulleriella viridis (Langerhans, 1881)2006TSALque
Chaetozone corona Berkeley & Berkeley, 19411982TSALest
Desdemona ornata Banse, 19571986TSALcas
Dispio magnus (Day, 1955)1982TSALcas
Dodecaceria capensis Day, 19611976TSALest
Dorvillea similis (Crossland, 1924)2014UNAALcas
Ficopomatus enigmaticus (Fauvel, 1923)1959TS-hullsALest
Glycinde bonhourei Gravier, 19042007UNAALcas
Hydroides brachyacantha Rioja, 19412015TS-hullsALcas
Hydroides dirampha Mörch, 18631981TS-hullsALest
Hydroides elegans (Haswell, 1883)1976TS-hullsALest
Leiocapitellides analis Hartmann-Schröder, 19602000UNAALcas
Leonnates persicus Wesenberg-Lund, 19492013UNAALcas
Lepidonotus tenuisetosus (Gravier, 1902)2008UNAALcas
Linopherus canariensis Langerhans, 18812007TCALcas
Lumbrinerides neogesae Miura, 19812002TSALcas
Lysidice collaris Grube, 18701975UNAALest
Mediomastus capensis Day, 19612006TSALcas
Neanthes agulhana (Day, 1963)2007TSALcas
Notomastus aberans Day, 19571964UNAALest
Paradyte cf. crinoidicola (Potts, 1910)1964UNAALcas
Polycirrus twisti Potts, 19281983UNAALest
Polydora cornuta Bosc, 18022008TSALcas
Prionospio pulchra Imajima, 19901991TCALcas
Protodorvillea biarticulata Day, 19631975TSALest
Pseudonereis anomala (Gravier, 1900)2003TSALest
Pseudopolydora paucibranchiata (Okuda, 1937)2005TSALest
Sigambra parva (Day, 1963)1975TSALest
Spirobranchus tetraceros (Schmarda, 1861)1970TS-hullsALest
Spirorbis marioni Caullery & Mesnil, 18971996TS-hullsALest
Timarete punctata (Grube, 1859)2006TSALcas
Bryozoa
Cheilostomatida
Celleporaria brunnea (Hincks, 1884)2015TS-hullsALunk
Celleporaria vermiformis (Waters, 1909)2015TS-hullsALest
Crepidacantha poissonii (Audouin, 1826)1986TS-hullsALunk
Crisularia serrata (Lamarck, 1816)1967TS-hullsALunk
Hippopodinasp. A as Hippopodina feegeensis (Busk, 1884)1996TS-hullsALest
Juxtacribrilina mutabilis (Ito, Onishi & Dick, 2015)2019-20TSALest?
Microporella coronata (Audouin, 1826)1967TS-hullsALque
Tricellaria inopinata d'Hondt & Occhipinti Ambrogi, 19852015TS-hullsALunk
Ctenostomatida
Amathia verticillata (delle Chiaje, 1822)1969TS-hullsALest
Crustacea
Cirripedia
Amphibalanus eburneus (Gould, 1841)1970TS-hullsALunk
Balanus trigonus Darwin, 18541970TS, TCALest
Megabalanus tintinnabulum (Linnaeus, 1758)1996TS-hullsALcas
Copepoda
Acartia (Acanthacartia) tonsa Dana, 18492005TSALest
Arietellus pavoninus (G. O. Sars, 1905)1967UNA, TSALest
Calanopia elliptica (Dana, 1846)1988UNAALest
Centropages furcatus (Dana, 1852)1988UNAALest
Euchaeta concinna Dana, 18491987ALest
Oithona davisae Ferrari F.D. & Orsi, 19842018UNA, TSALcas
Pseudodiaptomus marinus Sato, 19132016TSALest
Paracartia grani (G. O. Sars, 1904)1988TS-hullsALest
Parvocalanus crassirostris (Dahl, 1894)2009UNALest
Stomatopoda
Erugosquilla massavensis (Kossmann, 1880)1963UNAALest
Decapoda
Actaeodes tomentosus (H. Milne Edwards, 1834)2013UNAALcas
Alpheus rapacida (de Man, 1908)1998UNAALcas
Arcania brevifrons Chen, 19892021UNAALunk
Atergatis roseus (Rüppell, 1830)2009UNAALest
Calappa pelii Herklots, 18512005UNALcas
Callinectes sapidus Rathbun, 18961947TSALinv
Carupa tenuipes Dana, 18512009UNAALest
Charybdis (Charybdis) hellerii (A. Milne-Edwards, 1867)2004UNAALest
Charybdis (Goniohellenus) longicollis Leene, 19381996UNAALest
Coleusia signata (Paulson, 1875)2005UNAALest
Dyspanopeus sayi (Smith, 1969)2015TSALcas
Homarus americanus H. Milne Edwards, 18372019UNKALcas
Gonioinfradens giardi (Nobili, 1905)2010UNAALest
Ixa monodi Holthuis & Gottlieb, 19561999UNAALest
Matuta victor (Fabricius, 1781) 2018UNAALcas
Macrophthalmus indicus Davie, 20122009UNAALcas
Metapenaeopsis aegyptia Galil & Golani, 19901996UNAALest
Metapenaeopsis mogiensis consobrina (Nobili, 1904)1995UNAALest
Myra subgranulata Galil & Golani, 19902004UNAALest
Penaeus aztecus Ives, 18912012UNAALinv
Penaeus hathor (Burkenroad, 1959)2012UNAALest
Penaeus pulchricaudatus Stebbing, 19141995UNAALest
Portunus segnis (Forsskål, 1775)1991UNAALest
Thalamita poissonii (Audouin, 1826)1983UNAALest
Urocaridella pulchella Yokes & Galil, 20062018UNAALest
Trachysalambria palaestinensis (Steinitz, 1932)1995UNAALest
Xanthias lamarckii (H. Milne Edwards, 1834)2013UNAALcas
Amphipoda
Bemlos leptocheirus (Walker, 1909)2015UNAALunk
Caprella scaura Templeton, 18362002UNALest
Isopoda
Cymodoce fuscina Schotte & Kensley, 20052015TSALcas
Mesanthura cf. romulea Poore & Lew-Ton, 19862016TSALunk
Paracerceis sculpta (Holmes, 1904)2009TSALest
Paranthura japonica Richardson, 19092012TSALunk
Sphaeroma walkeri Stebbing, 19052015/2017TSALunk
Echinodermata
Ophiuroidea
Ophiactis savignyi (Müller & Troschel, 1842)1993UNAALcas
Echinoidea
Diadema setosum (Leske, 1778)2010UNAALest
Holothuroidea
Synaptula reciprocans (Forsskål, 1775)1995UNAALinv
Chordata, Tunicata
Ascidiella aspersa (Müller, 1776)1901UNALest
Ciona robusta Hoshino & Tokioka, 1967 (as Ciona intestinalis (Linnaeus, 1767))1901TS-hullsALest
Diplosoma listerianum (Milne Edwards, 1841)1996TS-hullsALest
Herdmania momus (Savigny, 1816)2010UNAALest
Microcosmus squamiger Michaelsen, 19272019TSALest
Polyclinum constellatum Savigny, 18162019TSALunk
Phallusia nigra Savigny, 18162008UNAALest
Styela plicata (Lesueur, 1823)1968TS-hullsALest
Symplegma brakenhielmi (Michaelsen, 1904)2015TS-hullsALest
Chordata, Pisces
Abudefduf sexfasciatus (Lacepède, 1801) 2017RELALcas
Abudefduf vaigiensis (Quoy & Gaimard, 1825) 2018UNALcas
Acanthurus cfr gahhm (Forsskål, 1775)2019RELALcas
Acanthurus sohal (Forsskål, 1775)2017CORALcas
Alepes djedaba (Forsskål, 1775)1960UNAALest
Apogonichthyoides pharaonis (Bellotti, 1874)1982UNAALest
Atherinomorus forskalii (Forster, 1801)1986UNAALest
Bregmaceros nectabanus Whitley, 19412014UNAALest
Callionymus filamentosus Valenciennes, 18372003UNAALest
Chaetodipterus faber (Broussonet, 1782)2019RELALest
Champsodon nudivittis (Ogilby, 1895)2012UNAALest
Cheilodipterus novemstriatus (Rüppell, 1838)2020UNAALest
Equulites klunzingeri (Steindachner, 1898)1946-64UNAALcas
Etrumeus golanii Di Battista, Randall & Bowen, 20122003UNAALest
Fistularia commersonii (Rüppell, 1835)2001UNAALinv
Fistularia petimba Lacepède, 18032021UNAALunk
Hemiramphus far (Forsskål, 1775)1943UNAALest
Lagocephalus guentheri Miranda Ribeiro, 19151952UNAALest
Lagocephalus sceleratus (Gmelin, 1788)2005UNAALinv
Lagocephalus suezensis Clark & Gohar, 19532003UNAALest
Lutjanus sebae (Cuvier, 1816)2010RELALcas
Lutjanus argentimaculatus (Forsskål, 1775)2019UNAALcas
Nemipterus randalli Russell, 19862018UNAALcas
Oncorhynchus kisutch (Walbaum, 1792)2021ALcas
Oxyurichthys petersi (Klunzinger, 1871)2010UNAALest
Pagrus major (Temminck & Schlegel, 1843)2019ECALest
Parexocoetus mento (Valenciennes, 1846)1946-64UNAALest
Parupeneus forsskali (Fourmanoir & Guézé, 1976)2017UNAALest
Pempheris rhomboidea Kossmann & Rauber, 18771983UNAALest
Petroscirtes ancylodon Rüppell, 18382004UNAALest
Planiliza haematocheila (Temminck & Schlegel, 1845)1995UNAALest
Pomacanthus imperator (Bloch, 1787)2016UNAALcas
Pomadasys stridens (Forsskål, 1775) 2019UNAALcas
Pteragogus trispilus Randall, 20131992UNAALest
Pterois miles (Bennett, 1828)2009UNAALinv
Sargocentron rubrum (Forsskål, 1775)1947UNAALest
Saurida lessepsianus Russell, Golani & Tikochinski, 2015<1971UNAALest
Scarus ghobban Forsskål in Niebuhr, 17752014UNAALcas
Scomberomorus commerson Lacepède, 18002008UNAALest
Siganus luridus (Rüppell, 1829)1964UNAALinv
Siganus rivulatus Forsskål, 17751925UNAALinv
Sillago suezensis Golani, Fricke & Tikochinski, 2014 2018UNAALcas
Sphyraena chrysotaenia Klunzinger, 18841995UNAALest
Sphyraena flavicauda Rüppell, 18382003UNAALest
Stephanolepis diaspros Fraser-Brunner, 19401943UNAALest
Synchiropus sechellensis Regan, 19082014UNAALest
Terapon theraps Cuvier, 18292008UNALcas
Torquigener hypselogeneion ex T flavimaculosus Hardy & Randall, 19832006UNAALest
Tylerius spinosissimus (Regan, 1908)2004UNAALest
Upeneus moluccensis (Bleeker, 1855)1947UNAALest
Upeneus pori Ben-Tuvia & Golani, 19892003UNAALest
Vanderhorstia mertensi Klausewitz, 19742019UNAALcas
Chordata, Mammalia
Sousa plumbea (G. Cuvier, 1829)2017UNAALcas

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